prop-ja:reference
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- Govaerts, R., Ruhsam, M., Andersson, L., Robbrecht, E., Bridson, D., Davis, A., Schanzer, I. & Sonké, B. . World Checklist of Rubiaceae. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; https://wcsp.science.kew.org/ Retrieved 7 September 2022 (ja)
- POWO . "Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30032120-2 Retrieved 25 October 2022 (ja)
- Govaerts, R., Ruhsam, M., Andersson, L., Robbrecht, E., Bridson, D., Davis, A., Schanzer, I. & Sonké, B. . World Checklist of Rubiaceae. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; https://wcsp.science.kew.org/ Retrieved 7 September 2022 (ja)
- POWO . "Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30032120-2 Retrieved 25 October 2022 (ja)
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